There is a fundamental difference between headphone listening and any other listening experience. With headphones, whether of the on-the-ear or around-the-ear type (that's what the "circumaural" in the specs means), the auditory canal is more or less on the axis of the headphone's diaphragm. In all other listening, much of the sound enters the ear only after reflecting off the pinna (ear lobe) and head. This factor alone drastically affects the frequency response perceived by the ear/brain.
John R. Sank, in a paper in the April 1980 Journal…