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One of my six pair of speakers, Bose 901-VI don't have removable grills. I have heard no coloration or alteration to the sound with the grilles on, but I'm over 60!!
An argument could be made that most speakers are designed to sound best with their grilles intact, while others might reason that anything in front of a tweeter has got to be bad. Reader Gerald Neily wants to know if you prefer to listen with your speaker grilles on or off.
I have B&W Nautilus 802s for mains, HTM1 for center, and four KEFs for surrounds. I keep the grilles on for appearance and protection, and keep them on for home theater and casual listening. I remove them from the 802s only for "critical" stereo listening because the sound is a bit more open. Only the mids and woofers have grilles; the tweeters are always exposed.
The dome tweeters on my speakers are vulnerable, so I usually think twice, but the grille cloths frequently come off when I am doing some earnest listening. It is a nice treat to remove them and hear more open highs and an extra layer of detail.
Depends on the mood. When I had a 5.1 system I would take the grills of now and again because I thought they sound brighter...but I'm beginning to realize how psychoacoustic sound really is. Lately i've been leaving them on (plus it's a pain taking off the grills in a 9.2 system)
The only time I put the grilles on to my Sonus Faber Concertos is for protection when I'm vacuuming the carpet around them or if children are around etc. As per Stereophile's review of them a few years ago, the 6dB midband suckout the grilles cause robs music of a little air and detail, and accentuates the treble harshness that digital sources foist onto our ears. Having the grilles off gives the sound a better balance.
Some years ago I owned a pair of Acoustic Research AR-3As. While the speaker's were audibly excellent the grilles were rather nondescript. Searching for audio neutral replacements turned out to be quite an undertaking but, in the end, well worth the time invested. The new grilles were audibly transparent and asthetically pleasing.